I can see this: Religious quests should absolutely be a thing, but for Jonoans, I think they aught have something different. I realize the idea is that it will incentivize conflict between them and the kypiq, however, I would say it needs to be a step up. Rather than just "A kypiq" I would suggest it be "A moshkin." These are, after all, fierce warriors, far more fitting of the Jonoan Hunt.
Outside of that, I think that Brudvir could have a couple different ones-- the tree one, okay, sure. But also, perhaps "slay a wolf den, and raise the wolven children- allow the elders to pass onto the next life, and raise their young in their stead."
Kypiqs... would be far more difficult. One can't protect the entire biome, on their own, so you'd really have to do that as a group effort, which while fitting for the kypiq, would also prevent players from reaching them if they come in late, or aren't available when other kypiqs do that mission.
Perhaps, you could also have one like: Collect foods to get through the winter, and leave it in the temple" and that would not only allow you to get spiritual actualization, but also it would provide food for your community.